Amazed by Toua's unique prowess on the mound, veteran slugger Hiromichi Kojima artfully scouts the pitcher for his long unsuccessful team, the Saikyou Saitama Lycaons. On the streets of Okinawa, he uses nothing but his wits and a "fastball" peaking at a mere 134 kmph to somehow achieve 499 wins in the game of "One Outs," a simplified version of baseball between the pitcher and one batter. Toua Tokuchi is an athlete by profession, but a reckless gambler at heart. With only three months left to qualify, Joe must face off against opponents the likes of which he has never fought in order to meet the challenge of his rival. Having no name of his own, he takes on the moniker of "Joe" as he begins his climb from the very bottom of the ranked list of fighters. Taken out in a single round, Junk Dog is left with a challenge: "If you're serious about fighting me again, then fight your way up to me and my ring." Filled with overwhelming excitement and backed by the criminal syndicate responsible for his thrown matches, Junk Dog enters Megalonia: a world-spanning tournament that will decide the strongest Megalo Boxer of them all.
However, this all changes when the Megalo Box champion Yuuri enters his shabby ring under the guise of just another challenger. Despite the young man's brimming potential as a boxer, the illegal nature of his participation forces him to make a living off of throwing matches as dictated by his boss Gansaku Nanbu.
"To be quiet and do as you're told, that's the cowardly choice." These are the words of Junk Dog, an underground fighter of Megalo Box, an evolution of boxing that utilizes mechanical limbs known as Gear to enhance the speed and power of its users.
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